Redefinition and generic revision of the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with descriptions of six new genera. (Bulletin of the American Museum of Natural History, no. 382) González-Santillán, Edmundo.; Prendini, Lorenzo.
The endemic North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, is redefined and its component genera revised, based on a simultaneous phylogenetic analysis of 250 morphological characters and 4221 aligned DNA nucleotides from three mitochondrial and two nuclear gene markers. Tribe Stahnkeini Soleglad and Fet, 2006, is removed from Syntropinae. Tribe Paravaejovini Soleglad and Fet, 2008, and subtribe Thorelliina Soleglad and Fet, 2008, are abolished: Paravaejovini Soleglad and Fet, 2008 = Syntropinae Kraepelin, 1905, syn. nov.; Thorelliina Soleglad and Fet, 2008 = Syntropinae Kraepelin, 1905, syn. nov. Eleven genera, six newly described, are recognized within Syntropinae: Balsateres, gen. nov.; Chihuahuanus, gen. nov.; Kochius Soleglad and Fet, 2008; Konetontli, gen. nov.; Kuarapu Francke and Ponce-Saavedra, 2010; Maaykuyak, gen. nov.; Mesomexovis, gen. nov.; Paravaejovis Williams, 1980; Syntropis Kraepelin, 1900; Thorellius Soleglad and Fet, 2008; Vizcaino, gen. nov. Hoffmannius Soleglad and Fet, 2008, is abolished: Hoffmannius Soleglad and Fet, 2008 = Paravaejovis Williams, 1980, syn. nov. Lissovaejovis Ponce-Saavedra and Beutelspacher, 2001 [nomen nudum] = Paravaejovis Williams, 1980, syn. nov. Ten species, formerly placed in Hoffmannius, are transferred to Paravaejovis: Paravaejovis confusus (Stahnke, 1940), comb. nov.; Paravaejovis diazi (Williams, 1970), comb. nov.; Paravaejovis eusthenura (Wood, 1863), comb. nov.; Paravaejovis flavus (Banks, 1900), comb. nov. [nomen dubium]; Paravaejovis galbus (Williams, 1970), comb. nov.; Paravaejovis gravicaudus (Williams, 1970), comb. nov.; Paravaejovis hoffmanni (Williams, 1970), comb. nov.; Paravaejovis puritanus (Gertsch, 1958), comb. nov.; Paravaejovis spinigerus (Wood, 1863), comb. nov.; Paravaejovis waeringi (Williams, 1970), comb. nov. Paravaejovis schwenkmeyeri (Williams, 1970), comb. nov., is removed from synonymy. Four species, formerly placed in Kochius, are transferred to Chihuahuanus, gen. nov.: Chihuahuanus cazieri (Williams, 1968), comb. nov.; Chihuahuanus crassimanus (Pocock, 1898), comb. nov.; Chihuahuanus kovariki (Soleglad and Fet, 2008), comb. nov.; Chihuahuanus russelli (Williams, 1971), comb. nov. Four species, formerly placed in Kochius, Thorellius, or Vaejovis C.L. Koch, 1836, are transferred to Mesomexovis, gen. nov.: Mesomexovis atenango (Francke and González-Santillán, 2007), comb. nov.; Mesomexovis oaxaca (Santibáñez-López and Sissom, 2010), comb. nov.; Mesomexovis occidentalis (Hoffmann, 1931), comb. nov.; Mesomexovis subcristatus (Pocock, 1898), comb. nov. Mesomexovis variegatus (Pocock, 1898), comb. nov., is reinstated to its original rank as species. Four subspecies are newly elevated to species: Kochius barbatus (Williams, 1971), stat. nov.; Kochius cerralvensis (Williams, 1971), stat. nov.; Kochius villosus (Williams, 1971), stat. nov.; Mesomexovis spadix (Hoffmann, 1931), comb. et stat. nov. Three subspecies are synonymized: Vaejovis diazi transmontanus Williams, 1970 = Paravaejovis diazi (Williams, 1970), syn. nov.; Vaejovis bruneus loretoensis Williams, 1971 = Kochius villosus (Williams, 1971), syn. nov.; Vaejovis hoffmanni fuscus Williams, 1970 = Paravaejovis hoffmanni (Williams, 1970), syn. nov
Morphological analysis of montane scorpions of the genus Vaejovis (Scorpiones: Vaejovidae) in Arizona with revised diagnoses and description of a new species
Several scorpions of the genus Vaejovis in Arizona are restricted in range to mountain-top forests. These scorpions, informally referred to as the “vorhiesi complex” are very similar morphologically, but their geographic distribution has attracted the attention of several researchers, resulting in the description of a few new species in recent years. However, these species were described from small sample sizes and were diagnosed with questionable characters that were not sufficiently analyzed. This study evaluates the morphology of scorpions of the “vorhiesi complex” from seven regions in Arizona to verify the validity of the species and their accompanying diagnoses. Morphological characters examined include morphometrics, hemispermatophores, size and shape of subaculear tubercles of the telson vesicle, pectinal tooth counts, pedipalp chela denticle counts, metasomal setal counts, development of metasomal carinae, and tarsal spinule counts. New diagnoses are given for previously described species (V. vorhiesi Stahnke 1940, V. lapidicola Stahnke 1940, V. paysonensis Soleglad 1973, V. cashi Graham 2007 and V. deboerae Ayrey 2009), which are considered valid, based on the morphological evidence gathered. A new species ofVaejovis, V. electrum, is described from the Pinaleño Mountains in Arizona.
Wernerius inyoensis, an elusive new scorpion from the Inyo Mountains of California (Scorpiones, Vaejovidae)
A new scorpion species is described from the Inyo Mountains of California (USA). The presence of a strong subaculear spine, along with other characters, places the new species withinWernerius, an incredibly rare genus that until now consisted of only two species. Wernerius inyoensis sp. n. can be most easily distinguished from the other members of the genus by smaller adult size, femur and pedipalp dimensions, and differences in hemispermatophore morphology. Previous studies have suggested that the elusive nature of this genus may be attributed to low densities and sporadic surface activity. Herein, we provide another hypothesis, that Wernerius are primarily subterranean. Mitochondrial sequence data are provided for the holotype.
The vorhiesi group of Vaejovis C.L. Koch, 1836 (Scorpiones: Vaejovidae), in Arizona, with description of a new species from the Hualapai Mountains
A new species in the vorhiesi group of Vaejovis C.L. Koch, 1836 (Vaejovidae Thorell, 1876), which appears to be endemic to the Hualapai Mountains near Kingman, Arizona, is described and illustrated. Vaejovis tenuipalpus, n. sp., the 11th species in the vorhiesi group, is compared to morphologically similar species, including V. jonesi Stahnke, 1940, V. lapidicola Stahnke, 1940, V. vorhiesi Stahnke, 1940, and V. deboerae Ayrey, 2009. The new species possesses the most slender pedipalp chelae in the vorhiesi group. New distribution records and a comprehensive distribution map are provided for all Arizona members of the group.
A new scorpion species, Vaejovis halli sp. nov., is described. This relatively small, brown new species is found on Mount Ord in the Mazatzal Mountains along the Mogollon Rim of northern Arizona. The new species appears most similar to V. vorhiesi Stahnke and V. deboerae Ayrey. The most distinguishing characteristic of this new species is the number of inner denticles (ID) found on the pedipalp fingers with six on the movable finger and usually five on the fixed finger, which more closely correlates with Vaejovis species from the mountains of southern Arizona rather than those geographically closer in northern Arizona.
Multivariate methods support the distinction of a new highland Vaejovis (Scorpiones: Vaejovidae) from the Sierra de los Ajos, Mexico
Multivariate analyses of morphological characters provide strong evidence that a highland Vaejovis from the Sierra de los Ajos, a Madrean ‘sky island’ in northern Sonora, Mexico, represents a distinct new species of the V. vorhiesi group. This new species is described and compared to other geographically adjacent species of the V. vorhiesi group, named
V. bandido, and brief notes on ecology are provided. Results from this study provide evidence that multivariate analysis of morphological characters is a powerful tool to delimit small and otherwise cryptic scorpion species.
A new Vaejovis C.L. Koch, 1836, the second known vorhiesi group species from the Santa Catalina Mountains of Arizona (Scorpiones, Vaejovidae)
A new species of the vorhiesi group of Vaejovis C.L. Koch, 1836, Vaejovis brysoni sp. n., is described from the Santa Catalina Mountains in southern Arizona. Vaejovis deboerae Ayrey also inhabits this mountain range, making this the first documented case of two vorhiesi group species distributed on the same mountain. When compared to all other vorhiesi group species, Vaejovis brysoni sp. n. is distinct based on several combinations of morphological characters and morphometric ratios.
Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City, ” Nevada (Scorpiones, Vaejovidae)
A new scorpion species is described from the Spring Mountain Range near Las Vegas, Nevada. The new species appears to be geographically isolated from other closely related species ofUroctonites Williams & Savaryand Pseudouroctonus Stahnke. We tentatively place the new species in Pseudouroctonus and provide detailed descriptions and illustrations of type material. We compare the new species to 17 congeneric taxa, briefly discuss the taxonomic history ofPseudouroctonus, and provide DNA barcodes for two paratypes to assist ongoing research on the systematics of family Vaejovidae.
Ayrey, R.F. 2013. A new species of Vaejovis from the Mogollon Rim of northern Arizona (Scorpiones: Vaejovidae). Euscorpius, 176: 1−13. (13 November 2013).
A new scorpion species, Vaejovis trinityae sp. nov. is described. This small brown species is found along the Mogollon Rim above Strawberry, Arizona. This is the first description of a new species of the “vorhiesi” group scorpions whose DNA phylogenetic analysis was published (Bryson et al., 2013); based on DNA data, the new species is most related to V. lapidicola Stahnke and V. crumpi Ayrey et Soleglad. It represents one of the “twenty-seven geographically cohesive lineages inferred from the mtDNA tree”. A unique characteristic of this species is that it exhibits arboreal behavior, being frequently found on Ponderosa pine trees.
Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones, Diplocentridae), with descriptions of three new species from Oaxaca, Mexico. (American Museum novitates, no. 3777)
The scorpion genus Diplocentrus Peters, 1861, comprising more than 50 species, most of
The scorpion genus Diplocentrus Peters, 1861, comprising more than 50 species, most of which are endemic to Mexico, is the most diverse in the family Diplocentridae Karsch, 1880 (Santibáñez-López et al., 2011). Hoffmann (1931) divided the Mexican species into two groups, the whitei group and the keyserlingi group, based largely on differences in size and coloration. Francke (1977) redefined these groups. The whitei group, renamed the mexicanus group because it included the type species of the genus, comprised species with short cheliceral fingers and the pedipalp femur wider than high. The keyserlingii group comprised species with long cheliceral fingers and the pedipalp femur higher than wide. Several new species of Diplocentrus were since described, but no attempt was made to synthesize the taxonomy of the species assigned to either group or further clarify the validity of the groups. In the present contribution, the species of Diplocentrus with the pedipalp femur higher than wide are reviewed. An operational diagnosis is provided for the keyserlingii group. Diplocentrus formosus Armas and Martín-Frías, 2003, previously synonymized with Diplocentrus tehuano Francke, 1977, is reinstated. Revised, updated diagnoses are provided for all previously described species and three new species, Diplocentrus kraepelini, n. sp., Diplocentrus sagittipalpus, n. sp., and Diplocentrus sissomi, n. sp., are described. The female of Diplocentrus mitlae Francke, 1977, is described for the first time. A dichotomous key is provided for identification of the 10 species in the keyserlingii group.